Bizarre yet endearing: Brave the world of mutant succulents
Crested and monstrose succulents — mutant forms only a succulent freak could love? We think not, whether it’s Halloween season or, say, Arbor Day…although these living oddities would be naturals in any décor scheme aimed at enticing or spooking trick-or-treaters and their parental units.
Euphorbia lactea cristata, also known as coral cactus
To start, the occurrence of growth mutations is documented throughout the plant kingdom, but we happen to be most interested in the topic as it applies to succulents, especially cacti and euphorbias. Mutations, however, occur in aeoniums, echeverias, and other succulents too.
Crested and monstrose plants are those that exhibit abnormal, distorted or disfigured growth. This happens when a mutation occurs during cell division. Crests are the elongation of the growing point from a radial one to a linear one. But rather than perfectly flat layers of tissue resulting, the damage to the central growing point creates multiple growing points that crowd and push against each other. Peculiar shapes form from this energy — such as those resembling brains, alluvial fans, coral, and snakes.
Cereus forbesii ‘Ming Thing’
In monstrose growth, the mutation typically makes random multiple growing points that are not stacked next to each other — there’s one here, another over there, pushing the body of the plant all out of shape in disorderly fashion. In monstrose plants, the mutation can manifest in a variety of freaky growth, from spiraling to extra ribs to the absence of spines or teeth, or even excessive teeth, like on an Agave ‘Cubic’ (Agave potatorum f. monstrose).
The crested and monstrose forms generally occur because of injuries to the plant at a young age. This damage can result from insects munching on the growing tip, or from other causes, including disease and genetic predisposition. In reaction to this injury, the cells at the tip of the branch where growth occurs begin to multiply at a much faster rate and the normal growing tip “goes crazy,” creating, in the case of crested plants, those fantastical fans and whorls.
Monstrose form of Myrtillocactus geometrizans
In monstrose specimens, each of those growth tips behaves as if it were the primary point. The result is messy, lumpy, monster-like growth. Cristates look more neat and symmetrical in comparison.
Plants can exhibit mutated and regular growth at the same time. Crests and monstrose growth are unpredictable; no two are precisely alike. That goes for plants of the same species. While we are not going to dive into the topic of cultivation here, those wanting to propagate their ornamental mutants can attempt to do so via cuttings, particularly those that include the abnormal growth. Propagation steps do differ a bit with these plants. The author of this article recommends that cuttings of crested cacti be grafted for best results. Lots of different examples of mutated succulents can be viewed in this piece.
Is that a curled-up snake?
So whether you incorporate your little monsters into your Halloween celebrations or not, we trust you love and cherish them just the same as you do your normal, “non-rule-breaking” succulents. These lovable freaks add to the amazing variety of succulent plants available to gardeners and collectors. It’s conceivable that one could possess, in addition to a beautiful, non-mutant specimen, a monstrose form of it, a crested version, and — why not — a variegated cristate type too.
Below are examples of monstrose and crested forms. Our online retail shop regularly harbors monsters and cristates. For online wholesale customers, we offer trays of assorted crested cacti and assorted monster cacti at The Cactus Shop.
Euphorbia flanaganii cristata ‘Green Coral’: Native to South Africa, this is one of the “medusoids,” or plants forming a central basal caudex with arms arising from the basal area. This is the cristate form of E. flanaganii, which forms deep emerald green fan-shaped stems that resemble green coral. Retail link.
Mammillaria gracilis v. fragilis monstrose ‘Arizona Snowcap’: A monster cultivar of the charming miniature Mammillariagracilis fragilis. It features thimble-shaped bodies to 1.5 inches in height with tufts of snow-white spines and wool at the areoles. Satiny, creamy yellow flowers come in late winter. Retail link.
Mammillaria elongata cristata ‘Copper King’: It’s known as the brain cactus because of its unusual growth, giving the appearance of, yeah, brains. Very attractive, with undulating fans covered in dense coppery orange spines. Retail link.
Opuntia microdasys monstrose
Opuntia microdasys monstrose: This paddle cactus is known as crazy bunny ears, an undulated form of regular ol’ bunny ears. It’s covered with minute golden spines known as glochids, which can cause irritation to the skin and eyes. Retail link.
Opuntia (Austrocylindropuntia) vestita cristata: Native to Bolivia, this cactus forms rounded stems in clusters that are densely covered with beautiful white hair. Vivid magenta flowers to 1 inch in diameter. Retail link.
Among the many (many) reasons we succulent lovers are fascinated by these plants is the extraordinary richness and diversity of forms and textures they display. We often come to prize the more unusual varieties. Nowhere, will you find more strange, unusual and incredibly cool forms than among crested succulents and monstrose cactus. Like most variegated succulents, both cristata succulents and “monster cactus” are the result of abnormal development of the plant. This is a naturally occurring phenomenon that is tricky to reproduce. Cresting causes wildly curved, twisted and undulating plants that look very different from un-crested forms. Crested cactus often look like writhing snakes, coral or even human brains. Maybe you’re intrigued by these curiosities. Or maybe you have a plant displaying strange growth and want to understand it. Learn why some succulents and cactus crest and how to care for them.
Crested Succulents & Monster Cactus
In this Post We'll Cover:
What are Crested Succulents and Cactus?
To understand crested succulents and cactus, let’s first look at how normal plants grow and develop. We’ve covered the nature of meristemMeristem (MEHR-i-stem) tissue in plants contains undifferent... tissue when we looked at propagating succulents by leaves as well as stem cuttings. Meristem tissue is made up of immature cells that can develop to become anything the plant needs – stem, root or leaf. Its like stem cells in animals.
The meristem tissue at the very tips of roots and shoots is called apical meristem. It forms a small point at the apex that is responsible for the plant’s primary growth. Apical meristem tissue makes the plant grow taller, developing more leaves for photosynthesisPhotosynthesis (FO-to-SIN-thuh-sis) is the process plants us... More. At the same time, apical meristem on the roots grow longer, wider-spreading roots to reach and access more water in the soil. These are critical functions for the plant. While all meristem tissue is capable of growth in the right circumstances, apical meristem tissue does grow and is responsible for the plant’s shape and size.
A crested cactus or succulent is the result of a genetic defect in the apical meristem. The apical meristem tissue of a normal saguaro cactus cause the classic vertical, columnar shapes. Occasionally the cactus branches, but most of its growth continues to reach for the sky. When a saguaro crests, the apical meristem no longer forms a single point. Instead, it forms a broad line, with new growth points forming all along that line. The resulting plant shape becomes wildly distorted and contorted along that line. As the plant grows taller and wider along the resulting crest, the plant begins to curl and fold in upon itself. A crested succulent or cactus has the Latin word cristata added to its botanical name. So we call a saguaro (Carnegiea gigantea) that crests a Carnegiea gigantea cristata.
What is a Monstrose Cactus or Succulent?
Monstrose is another form of a genetic defect in the apical meristem of a plant. The organized points of new growth become random, forming all over the body of the plant, causing new growth to form anywhere. A crested cactus or succulent still displays organized growth with a predisposition toward symmetrical development. A monstrose cactus, however, develops in random growth points all over the plant. This results in a lumpy, knobby, asymmetrical shape. Sometimes monstrose cactus develop bizarre corkscrew spirals or extra ribs with very few or deformed spines. Like crested succulents, monstrose cactus are highly sought after and collectible plants.
Monstrose cactus are often referred to as monster cactus. The two words share the same root word — monstrum, meaning birth defect with abnormal, disfigured growth. While animals with weird deformities might appear to us as monsters, in plants, the same genetic mix-ups often turn out looking bizarrely cool.
What Causes Cresting and Monstrose Development in Succulents?
Although both cresting and monstrose forms are the result of genetic defects, they cannot be inherited. Generally, a crested or monstrose plant is the result of an injury to the apical meristem of a young plant. This injury might be the result of wind, weather, insects, bird or animals. Sometimes a nutritional deficiency or hormone imbalance can lead to cresting or monstrose. It can result from disease or infections, or even seem to occur spontaneously when cells divide improperly. When the cells respond to the injury with fast, somewhat organized growth, it forms a crest. When the new growth occurs all over the plant, that is a monstrose. Any type of plant can develop a crest or monstrose, but they do seem more common with cactus and succulents. Likely, the tough nature of these plants makes them better able to survive to maturity even with genetic defects than other plants.
Because cresting and monstrose formations result from injury to the plant, the abnormal growth is not typically passed on to the plant’s offspring. An injured parent doesn’t develop an injured offset. However, some cactus and euphorbia varieties are more likely to crest than others. And this predisposition is hereditary. You may see both crested and normal development on the same plant. You may even see all three forms, with monstrose growth in the mix. A plant may crest at any point in its life, but monstrose typically develop only in very young plants.
There are a very few varieties that do reliably pass their peculiarities on to their offspring and can be readily reproduced through cuttings. Notable among these are the Crassula ovata Hobbit, Gollum and ET’s Fingers varieties of the ever-popular jade plant. Not only do these charming plants resemble the outstretched fingers of some inhabitants of Middle Earth, but these succulents also thrive in low light, indoor conditions.
Care for Crested and Monstrose Succulents
Every crested and monstrose succulent develops into a fantastical form unique to the individual plant. Plants evolve their growth rates, shape and sizes over millions of years to enable them to thrive in their native habitat. So it’s no surprise that these one-of-a-kind plants are a bit more sensitive to environmental pressures than their un-crested counterparts. Crested and monstrose cactus and succulents are actually quite stable and able to grow for years with care. They tend to be even more sensitive to over-watering and fertilizing than un-crested plants of the same species. The abnormal tissues are more prone to splitting, rotting or additional injury. Provide the same lighting and temperature controls you would for that succulent variety, just give the crested form even less water, less often and with less fertilizer. This is known as growing the plant “harder”.
PropagatingPropagating (PRAH-puh-gate-ing) a plant is to grow another p... and Pruning Crested and Monstrose Cactus
Because crested and monstrose succulents are the result of injury and not heredity, the fabulous forms are not passed down to the plant’s offspring. You cannot get more crested succulents through collecting and sowing seeds from the plant. Instead, crested succulents are propagated from cuttings taken from a part of the plant that is cresting. These cuttings are generally grafted to a hardy cactus rootstock, the way moon cactus are. Grafting helps to make the plant more stable, as the root structure is provided by a vigorous, un-injured plant.
In time, you may see a normal succulent develop a crest. Sometimes a crested or monstrose cactus develops normal growth. It may sprout from the rootstock or the plant may revert to normal growth patterns. If you want to maintain the crested form, cut back any shoots of normal plant growth. In time, normal growth can out-compete the crested growth and cause the entire plant to revert.
Buying Crested Succulents and Monstrose Cactus
Many succulent lovers go wild for the bizarre forms of crested and monstrose succulents. Although they are rare, you can find good quality crested succulents pretty easily. Sometimes, even at big box stores. My favorite succulent vendors are great sources for these spectacular plants. Leaf & Clay offers an intriguing array of 16 different named monstrose and crested succulents. For some, like Echeveria ‘Cubic Frost™’, they carry both the normal and crested varieties. Mountain Crest Gardens offers a few crested succulents as well as a few monstrose varieties. The Succulent Source has an ever-changing selection of crested and monstrose succulents and cactus. Some of their selection, especially the larger specimens, allow you to choose the exact crested or monstrose plant you want.
You’ll also find crested and monstrose succulents on Amazon and here on Etsy. Wherever you get your plant, I hope you’ll give one of these phantasmagorical beauties a try!
I hope you’ve enjoyed this walk on the wild side. Do you already grow crested succulents, or do you plan to give one of these phantasmagorical beauties a try? I’d love to hear from you! Please take a moment to leave me a comment and let me know. Especially if you have any questions!
’til next time,
P.S. For more succulent info — and to just make my day! — please subscribe to The Succulent Eclectic! I’ll send you my FREE course, 7 Steps to Succulent Success. Thanks!
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Cristate and Monster Succulents
Succulents (including cacti) are among the best of all the plants to collect and grow in pots and in the garden and are certainly among my favorite plants.As if there weren't enough awesome, cool and bizarre succulents in cultivation to keep one fascinated and obsessed for a few lifetimes, these amazing plants also have their mutant forms- the crested and monstrose plants.Crested or monstrose mutations are not unique to succulents, but they certainly seem to be much more common in these plants than they are in most other flowering plants and conifers.
What happens when a plant mutates like this?
All plants have a growth center (or growth centers depending on how many branches, leaves, flowers etc. they have).On most succulents that have been found to crest, this apical meristem is where the primary growth of the plant occurs and normally these primary growth centers produce cells from a single point and as new cells are formed all the older cells are moved outward in a nearly radially symmetrical pattern (creating columns or branches that tend to be circular in cross section).
This Sempervivum was one of hundreds I have struggled to grow in the garden over the years (most are pretty wimpy in this climate). But for some unknown reason, this particular plant began to crest. You can almost see the linear plane of tissue growing out each side, when it SHOULD be growing from a central location radially.
These two plants also spontaneously crested in my garden, though the crested areas are harder to see since they are producing lots of dinky leaves (relatvie to the normal leaves). Aeonium 'Sunburst' (a variegate- another mutation seen frequently in succulents) on the left and Echeveria elegans on the right.
But when a mutation occurs in this apical meristem, often a linear row of meristematic cells develop creating a growth pattern that originates from a line rather than a single point.This mutation is what causes cresting (aka fasciation) to occur and flattened planes of new growth to develop.Instead of forming perfectly flat sheets of tissue more often curling develops as a result of external and natural internal forces. This results in the formation of ornamental waves or walls of new growth that can develop into a wide variety of interesting shapes and patterns (in cacti, many form 'brain-like' structures).Crested plants often look very different than their normal counterparts, but usually there is some recognizable tissue and/or coloration, and often normal growth areas grow out of the same plant alongside the crested areas. Crested plants can develop their mutation early in life but more often a crest develops later on, so a normal looking plant will develop a crested section. Plants that crest early in life tend to stay somewhat immature and rarely flower or develop a reproductive functional anatomy. Those that creat later can usually flower and reproduce normally.
Here are two Aeonium arboreum 'Atropurpureum' crests. It looks like the one on the left crested earlier in life (though this could be a rooted cutting) since the lowest portion of stem is flattened (when it should be cylindrical)... but the top almost looks like it's 'defasciating' or reverting. Right plant is a mature individual that developed a crest mutation on one of its branches. These two crested individuals show a flattened plane of growth
These two cacti, on the other hand, show more typical undulation and curving that occurs as crests continue to develop and mature. The Parodia densispina on the left is showing the typical 'brain'-like appearance of many cactus crests. The Ariocarpus retusa on the right resembles a tidal sea creature thanks to its gorgeous undulations mixed with its more typical tubercular shape. Both these cacti retain some of the plant's original coloration and spination (or lack thereof), but do not really look anything like the normal plants.
These Euphorbia lactea crests show the more typical fan-like undulation seen in most cristate Euphorbia species. Some might notice this is the same photo in the other mutant plant article of the month- the one on variegated plants, as many of these crested Euphorbias also show varying degrees of variegation as well. Notice these plants are all grafted on a different Euphorbia species- one with a healthy root stock and probably one much more able to photosynthesize and deal with more irregular watering schedules.
These two cacti are NOT crested, though many think these are crested plants. These are two of many species of cacti that develop special areas, either on the top (as in the Melocactus species on the left) or, more commonly, along the top and down one side (like the Coleocephalocereus aureus cacti in the right photo) called cephalums. Cephalums are not mutations, however, and are a genetically consistent structure these species form with age and where the flower parts arise from.
Pachypodium lamerei crests are fairly common mutations, but they eventually outgrow themselves and collapse
Crests to do not always form in the same tissues, and where they form will create a different look even in the same species. Left is a normal form of Euphorbia flanaganii, a medusoid Euphorbia frequently found to crest. Right is a very common crested form of this species in which the 'arms' of the medusa have all (or almost all... you can see a few normal arms at the bottom of the photo and mixed in here and there) crested. This form of cresting is so common that these crested Euphorbia flanaganiis are frequently encountered at garden outlet shops.
This specimen above was one I planted in my yard. It was completely normal at the time, but grew rapidly thanks to my overwatering and the very rich, fertile soil. And sure enough, a crest formed in my plant.
The Euphorbia flanaganii on the left has a cristate mutation in the tissue that forms at the top of the stem from which the 'arms' arise. Therefore this plant has a linear, curving ridge of crested basal tissue but normal looking arms (sort of... most of the arms are a bit stunted and irregular). The much rarer form of Euphorbia flanaganii crest on the right is showing a cristate or fasciate mutation of the central stem of the medusoid structure and only somewhat involving the basal 'green' tissues.
One of the more spectacular crested plants I have seen photos of is this amazing Agave geminiflora crest of a flower stalk on the right (photo by Xenomorf); a normal flowering plant is shown in the lefthand photo. I think this is the only cristate flower stalk I have seen in a succulent species.
When mutations occur in a more systemic fashion (rather than at a single growth point) involving all the meristematic tissue in a plant, the result is what is known as a monstrose pattern of growth.In these plants all or most of the growing points start to develop haphazardly (sort of like how cancerous tumor grows in our bodies) and the result is a lot of knobs, bumps, asymmetrical and shortened limbs, branches or columns and generally a weird, stunted and freakish overall appearance.These monstrose plants are often so different from the unaffected plants that the two do not look related.In my limited experience, most monstrose plants rarely have normal plant parts growing alongside the weird ones, but it does happen now and then. Monstrose plants tend to develop their mutation early on rather than later in life.
left is a crested form of Astrophytum myriostigma, a commonly grown spineless cactus available at most succulent nurseries. The crested plants are quite rare however (photo cactus_lover); on the right is a Monstrose form of this same species. So instead of the plant growing along a linear plane, now numerous areas of abnormal mutant growth occur all over the plant. This form has been called Astrophytum myriostigma 'Lotusland' as that is where this particular clones was propagated.
The above Cereus cacti (Cereus jamacara left and Cereus hildmannianus right) are typical monstrose looks for many cacti, but particularly the Cereus genus.
The large Cereus species growing outdoors in the Huntington Gardens is a monstrose individual that is also showing a lot of cristate mutations as well
Myrtilocactus geometrizans is a very common species to see cresting (left), but pretty rare to see with a monstrose mutation (right- photo Kada)
The monstrose form of Mammillaria bocasana (normal form in left photo) is so unlike its normal version that for years these monstrose mutations were thought to be a completely different cactus species. But then some would show a bit of a reversion (defasciation) as seen in the plant on the right. This monstrose Mammillaria bocasana has been sold and grown in the nursery field as Mammillaria 'Fred'. It is one of the more popular and sought-after monstrose cacti
Both the giant, 20-foot-tall cactus on the left (Cereus hildmannianus subsp. uraguayanus- photo BayAreaTropics) and the 12-inch-tall mature plant on the right are the same subspecies of cactus. Only the plant on the right is a monstrose version of the monster on the left called 'Fairy Castle', and is a very popular and commonly available monstrose cactus in cultivation.
Pachycereus (aka Lophocereus) schottii is an example of a monstrose mutation seen in several other species of cacti in which the plants have lost some to all their spines and all the edges are more rounded. These monstrose forms are so common and popular that finding the original form (left photo) are so rare, some think that all Pachycereus schottii are lumpy, spineless plants. There are two distinct forms of this monstrose cactus- a fat one and a skinny one. This plant on the right in my yard is one of the fatter ones. These are excellent landscaping and potted plants- very hardy and easy to grow (from cuttings, too) and other than their relatively shorter stature and much slower growth, are excellently adapted plants to a variety of enviroments.
In Agaves, the monstrose mutation takes on a completely different look, basically stunting and somewhat deforming leaves, but not making the plant look all that different from its original shape and color. Agave 'Cornelius' on the left is a monstrose form of Agave americana variegata and grows much more slowly, is much smaller and has slight twisted stunted leaves... but it is a great, hardy potted or landscape plant. Right is a rare monstrose form of Agave titanota showing stunted, somewhat fused leaves and multiple growth centers.
Cycad monstrose mutations are generally limited to the leaf meristem and as a result one gets repeated monstrose changes in each leaf resulting in irregularly deformed leaves, often thicker than normal and usually split rather than smooth-edged and entire. Left is Ceratozamia hildae monstrose, and right is Zamia splendens monstrose
Why does this happen?
The precise cause of these mutations is unknown.Many suppose a bacterial, fungal or viral infection may cause some genetic disturbance.Another pathogen called a phytoplasma (sort of half way between a bacteria and a virus) has been shown to be the causative agent of cresting in at least some species.Chemical and physical trauma have to be included in the list of possibilities, though usually that sort of occurence damages the meristem (growth center) in such a way that it simply begins to divide, resulting in 'ordinary' branching or multiple heads. But for some reason, sometimes the result is crestation instead.I am sure pesticides and other toxic chemicals can alter gene function and result in mutations like this as well.Perhaps radiation from the sun is another possible mutation cause.Some plants seem more prone to this mutation at various seasons, so temperature, humidity or heat may have some influence.Some nutritional deficiencies have been known to lead to cresting mutations (e.g., Zinc deficiency) as well. And sometimes mutations can occur spontaneously (no external force needed)- just a chromosomal aberration resulting from some internal error in transcription or translation.
Either way, it has been difficult to nearly impossible to artificially produce many crested or monstrose mutations in cultivation.Such mutations are rarely heritable (so seed from such plants only produce more normal plants), at least in succulents; some dicots have been shown to have a heritable fasciation gene. But there has to be some sort of genetic tendency for these mutations to occur as many species form crests fairly commonly while others have never been found to do so.It is interesting to note the most succulents and cacti that form crests are those designed for extreme environments, while those who have evolved in more humid, temperate climates rarely crest (some exceptions of course).And there are some cacti and Euphorbia species where crested individuals are so common that one can obtain them at local nurseries as cheaply as the normal plants. Some are so common that it will not be unusual to so some purchased, cultivated normal plants to start cresting spontaneously in the garden. There still is a lot of research that should be done in this field.
Do crested and monstrose plants grow slower than normal ones?
Yes and no.Initially it appears that many crested species grow faster which is not surprising as there are many more growth points.But this increased rate of tissue production seems to have its limits, and when the plant tissues begin to become crowded, they often seem to cease growth, and often even form isolated spots of ‘defasciation' (reversion to normal growth), and several columns of normal plant material will erupt from this stalled crest formation.Most growers of prized crested plants will quickly remove these areas of reversion so the plant will concentrate its efforts on keeping the crested tissues happy and alive (and cool looking).
Both Crassula ovata 'Hobbit' and Crassula ovata 'Gollum' are mutations of the original species, though Crassula 'Hobbit's leaves are only slightly affected while those of 'Gollum' are truly monstrose. The two plants in the photo on the left are the same age, yet the 'Hobbit' on the far left is nearly twice the size as the 'Gollum' of the same age. Right is an old Euphorbia ingens monstrose form... about three feet tall... yet a normal Euphorbia ingens that age would probably be closer to 30 feet tall by now.
This Euphorbia lactea 'Ghost' crest (left) is showing reversion of the cristate growth back to normal, upright growth (defasciation). Most collectors would cut off all the new normal growth to keep the bizzare look intact, but this plant is sitting in the back of a nursery. Right is a weird monstrose Mammillaria that is growing some normal, defasciated tissue seen near the back right side
Monstrose plants on the other hand definitely seem to grow more slowly from day one.These plants are often stunted, though not all plants are monstrose 'all over'. Some have selected areas of monstrosity (eg. leaves only) and these plants tend to be more normal in shape and size.
Are crested and monstrose succulents more sensitive and harder to keep alive?
In general, these plants are more sensitive and touchy, which is really not surprising.Plants have evolved to grow in certain rates, shapes and sizes for a reason, and mutated plants that grow differently are very likely to be less well adapted to the original environments and may succumb.Growers of crested and monstrose plants often find they have to grow their plants a bit ‘harder' (water even less than normal) as the abnormal tissues are more likely to crack, grow too fast in spots and become injured, or simply rot, due to their physical changes from the ordinary.Fortunately plants in cultivation are not at the same ‘environmental mercy' as plants in the wild are, and probably far more crested plants survive in cultivation than in the wild (depends on the grower of course... plants under my care are really subject to abuse that even the wild could not match... as a result, I have few surviving crested or monstrose plants in my collection.Those that survive year after year are extremely tolerant, hardy plants!)
This plant (Echinopsis species?) was beautiful... until it rotted where the heavy cristate segments sat in the soil- oops.
This is one of the reasons crested succulents grown in cultivation are grafted onto other plants- often very hardy and non-fastidious species with relatively minor nutritional needs.These grafted plants rely upon their base species to absorb water and perform many of the biochemical pathways needed for proper growth that they do not perform as well.But these basal plants often continue their own struggles to not only survive, but become dominant, and healthy new growth will often branch off them.These branches need to be removed in a timely fashion or the mutational graft may suffer..
These crested cacti for sale in the left photo are all grafted plants- less like to die, and more likely to stay healthy that way; Euphorbia piscidermis crest on right hand photo grown on a Euphorbia canariensis cutting
Probably most monstrose mutations die early on, possibly so early they do not even develop far enough for one to recognize they are monstrose. So it is only the monstrose mutations that are compatible with life that we get to see grow far enough for us to recognize them and collect and marvel at them. Who knows what percentage of all monstrose mutations these survivors are?
Some crested and particularly some monstrose succulents, however, seem to be perfectly adapted and hardy, or at least not noticeably less so than their normal relatives.If the mutation does not affect the plant's ability to photosynthesize, grow upright and remain stable, and deal with water similarly, there is little reason such a mutation will affect the plant negatively.Though one wonders then why such mutations do not seem to take over in the wild and develop eventually into their own species.Perhaps, over time, they will?
This Crassula 'Gollum' though a bit slower and shorter than the normal Crassula ovata, is as easy to grow, as hardy and flowers as well as the normal plants in my garden
Below are just some of the crested and monstrose plants I have seen (and sometimes grown) in southern California at various cactus and succulent shows, nurseries or in various botanical or private gardens.
These are are show quality Aeonium cristate plant. Upper left Aeonium arboreum 'Atropurpureum', upper right was not labeled, but looks like Aeonium balsamiferum (hard to tell when crested), lower right is a rare species, Aeonium simsii.
Left is a crested form of Echeveria agavoides; right is a Senecio cylindricrus crest
Agave filifera and Agave titanota monstrose forms, also in Los Angeles cactus shows
Left is a grand prize winning Ariocarpus retusa crest; right, for reference, is a normal Ariocarpus retusa plant
Left and below are two crested Echinopsis species (lower might be Echinopsis pachinoi); right is a monstrose Austrocylindropuntia sublata (photo by Kell)
This crested plant has a few normal columns coming up in front
left is a crested Cereus species in Lotusland, Santa Barbara, California; right is a nice crested Cephalocereus urbanianus in the Huntington conservatory
left is a monstrose version of Cleistocactus strausii and rigth is a crested Cleistocactus winterii, both in my garden
left is a normal Copiapoa tenuissima while right is a commonly encountered monstrose form of the same species
left is a crested Copiapoa cinerea for sale at a local nursery (better hurry, this a rare one!); right a crested Discocactus zehntneri
Golden Barrel Cacti (Echinocactus grusonii) are fairly common landscaping plants, and reach enormous sizes, whether as singe, massive spheres, or large clumps (right). But this is also one of the more commonly available crested species of barrel cactus (see below)
Left is a more typical crest just starting out, for sale of Echinocactus grusonii; right is a much rarer and very bizzare crested form that again resembles a brain
The Lemon Ball Cactus (Parodia lenninghausii) is one of the more commonly available cacti for both landscape and pot use (left), but it looks very little like its mutant, crested form on the right
left is a normal Haageocereus pseudomelanostele cactus, and below is its monster equal, a crested plant; right is a monstrose version of Ferocactus cylindraceous (photo Xenomorf)obviously handling the stresses of nature about as well as non monstrose plants do
Haageocereus pseudomelanostele crest
These monsters also seem to survive in nature OK. Left is a cristate Carnegiea (Saguaro Cactus)- photo Franj; and right is a monstrose version of the same species (photo CactusJordi)
left are more of these Pachycereus schottii monstrose plants that are so popular, showing their typical gnarled, knobby, spineless shapes; right is another 'bald' form of monstrose cactus- Trichocereus (aka Echinopsis) bridgesii
left is a monstrose Opuntia basilaris (or maybe cristate?); right is one of my favorite of all the monstrose cacti, a Eulychnia sp. which grows like a tight spiral corkscrew and is one of the most bizzare and ornamental of all the cactus monsters in my opinion.
One of the most popular and well known mutant cacti is the Opuntia microdasys var. pallida monstrose form (normal form on left) called 'Funny Bunny'. This monstrose cactus I find grows very well but eventually gets too leggy and pieces fall off... though I am not sure that doesn't happen with the normal plants, too. It is a hardy mutant, however, and a good landscape plant. Some may say these are cristate, but Opuntia pads are already a fan shape, and what you see in the 'Funny Bunny' plants is bizzare and totally irregular growth patterns with folded pads, stunted shapes and all sorts of weird, irregular growth patterns, There is no real linearly bases cristate growth pattern, but its already fan-shaped pads make it look like this is a cristate mutant.
Though I have dozens and dozens more photos of cristate and monstrose cacti I could show here, I am rapidly running out of room. But no article on cristate cacti would be complete without a few shots of the original 'brain cactus', the cristate form of Mammillaria elongata (normal, though copper- colored, plant on right).
These tightly undulating globoid brains, complete with sulci, are very common cristate mutants of a pretty common garden Mammillaria. These are not particularly hard to keep alive, but they do rot easily if grown in the ground, so I put mine in a raku pot I made (right), where it has lived for years.Reportedly they reach this size fairly quickly, but then seem to simply stop growing (just continue to grow but very slowly)
This species also comes as a nice monstrose form, which also does very well in pots and I have had one of these for years, too, in the garden
a prize winning crested Euphorbia called Euphorbia 'Garretson' (or was labeled as such)
These Euphorbia lacteas are remarkably large for crests, which tend to eventually collapse on themselves. Plant on right is nearly a complete albino as well
Euphorbia susannae crest upper left, monstrose upper right and normal left
Euphorbia obesas make very interestig crests, like the plant on the left just starting to crest, the one on the right looking like a cactus 'brain' and the show winning plant at the top right of this article.
Euphorbia lambii, a common landscape tree/shrub in southern California, is one of the most commonly seen Euphorbias to crest. Just about every local botanical garden has at least one of these, and I have grown several that showed up spontaneously in my own garden (but eventually rotted). Normal looking plant lower left photo
Euphorbia millii (Crown of Thorns) crest (an exceptionall one)
Euphorbia millii crests for sale at a nursery (left); one of the most common Euphorbias to crest is Euphorbia nerifolia. Any many of these also are variegated, as is my plant (right). These are pretty hardy plants for crests and I have had no trouble keeping this one alive for years.
Luckhoffia beuneckeri is an Asclepid (flowers smell like rotting meat) and one of the most commonly encountered Asclepids to crest. This remarkable specimen was mine, but sadly I did not care for it well. Last only a few few years (not a good outdoor plant in my area).
A nice specimen in a plant show (still not as nice as the one I had for a while)
left is a normal Luckhoffia beuneckeri (photo Xenomorf); right is the only other cristate Asclepid I have seen, a Huernia piladdiii
Two last photos: Pedilanthes spontaneously cresting at the Los Angeles Zoo (left) and an unknown Yucca species cresting (and winning) at a show (right)
There are so many more oddities and curiosities out there, but I have already taken up far more space intended for these articles. For more about crests and other monsters see the following links:
Call Zoya, we will wash the purchase. Zoya turned out to be an extremely pleasant girl, as beautiful and intelligent as Masha and with the same mischievous and lustful eyes. She also burned a hole in me, most likely because of Masha's stories, which I am sure reached her.
Or daughter with his own mother does not inevitably lead to regression or disintegration of the personality of the incest. At the same time, they do not say this in relation to the sexual relationship between the daughter and the father, and the revealed relationship is regarded. As molestation of a minor child. This is a fact of life and the attitudes of society, which you need to adhere to and restrain your ardent sexual desires.
Despite the above, life makes its own adjustments and you need to understand that incest, like sex between a mother and a son or daughter, like sex between a.WEEKLY PLANT SHOP ROUTINE! - Running an Online Houseplant Shop from Home!
Cancer. Daddy took a glass dildo and some lubricant from the table, lubricated it and came up to me. - Since this is your first time, it will work out your ass first. When he put the glass head to the anus, I shuddered, the dildo was very cold. Putting my hands on my buttocks, I spread them apart and daddy began to slowly twist it.
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